Characteristics
Early sarcopterygians or crossopterygians are bony fish with fleshy, lobed, paired fins, which are joined to the body by a single bone. The fins of crossopterygians differ from those of all other fish in that each is borne on a fleshy, lobelike, scaly stalk extending from the body. Pectoral and pelvic fins have articulations resembling those of tetrapod limbs. These fins evolved into legs of the first tetrapod land vertebrates, amphibians. They also possess two dorsal fins with separate bases, as opposed to the single dorsal fin of actinopterygians (ray-finned fish). The braincase of sarcoptergygians primitively has a hinge line, but this is lost in tetrapods and lungfish. Many early sarcopterygians have a symmetrical tail. All sarcopterygians possess teeth covered with true enamel.
Taxonomists who subscribe to the cladistic approach include the grouping Tetrapoda within this group, which in turn consists of all species of four-limbed vertebrates. The fin-limbs of sarcopterygians such as the coelacanths show a strong similarity to the expected ancestral form of tetrapod limbs. The crossopterygians apparently followed two different lines of development and are accordingly separated into two subclasses, the Rhipidistia (including the Dipnoi, the lungfish, and the Tetrapodomorpha which include the Tetrapoda) and the Actinistia (coelacanths).
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