Tortoise - Taxonomy

Taxonomy

The following species list largely follows Rhodin et al., 2010, this is a work in progress.

Family Testudinidae Batsch 1788

Subfamily Testudininae

  • Aldabrachelys
    • Aldabrachelys gigantea, Aldabra giant tortoise, common synonyms Geochelone gigantea, Dipsochelys gigantea
    • Aldabrachelys abrupta (extinct)
  • Astrochelys Gray, 1873
    • Astrochelys radiata, radiated tortoise
    • Astrochelys yniphora, angulated tortoise, (Madagascan) plowshare tortoise, angonoka
  • Chelonoidis Fitzinger, 1835.
    • Chelonoidis carbonaria, red-footed tortoise
    • Chelonoidis chilensis, pampas tortoise, South Argentine tortoise
    • Chelonoidis denticulata, Brazilian giant tortoise, yellow-footed tortoise
    • Chelonoidis nigra complex:
      • Chelonoidis abingdonii, Pinta Island giant tortoise, Abingdon Island giant tortoise (extinct)
      • Chelonoidis becki, Volcan Wolf giant tortoise, Cape Berkeley giant tortoise
      • Chelonoidis chathamensis, San Cristobal giant tortoise, Chatham Island giant tortoise
      • Chelonoidis darwini, San Salvador giant tortoise, James Island giant tortoise
      • Chelonoidis duncanensis, Pinzon giant tortoise, Duncan Island giant tortoise
      • Chelonoidis hoodensis, Espanola giant tortoise, Hood Island giant tortoise
      • Chelonoidis nigra, Floreana giant tortoise, Charles Island giant tortoise (extinct)
      • Chelonoidis phantastica, Fernandina giant tortoise, Narborough Island giant tortoise (extinct)
      • Chelonoidis porteri, Santa Cruz giant tortoise, Indefatigable Island giant tortoise
      • Chelonoidis vicina, Isabela Island giant tortoise, Albemarle Island giant tortoise
    • Chelonoidis petersi, Chaco tortoise, northern Argentine tortoise
  • Chersina
    • Chersina angulata, Angulate Tortoise, South African bowsprit tortoise
  • Colossochelys
    • Colossochelys atlas, Atlas tortoise (extinct)
  • Cylindraspis (all species extinct) following Austin and Arnold, 2001:
    • Cylindraspis indica, synonym Cylindraspis borbonica, Reunion giant tortoise
    • Cylindraspis inepta, Mauritius giant domed tortoise
    • Cylindraspis peltastes, Rodrigues domed tortoise
    • Cylindraspis triserrata, Mauritius giant flat-shelled tortoise
    • Cylindraspis vosmaeri, Rodrigues giant saddleback tortoise
  • Geochelone
    • Geochelone elegans, Indian star tortoise
    • Geochelone platynota, Burmese star tortoise
    • Geochelone sulcata, African spurred tortoise (Sulcata tortoise)
  • Hadrianus
    • Hadrianus corsoni (syn. H. octonarius)
    • Hadrianus robustus
    • Hadrianus schucherti
    • Hadrianus utahensis
  • Homopus
    • Homopus areolatus, parrot-beaked tortoise, beaked Cape tortoise, common padloper
    • Homopus boulengeri, Karoo dwarf tortoise, Boulenger's Cape tortoise, Karoo padloper
    • Homopus femoralis, greater dwarf tortoise, greater padloper
    • Homopus signatus, speckled tortoise, speckled padloper, speckled padloper tortoise
    • Homopus solus, Nama tortoise, Berger's cape tortoise, Nama padloper, synonym Homopus bergeri
  • Indotestudo
    • Indotestudo elongata, elongated tortoise, yellow-headed tortoise
    • Indotestudo forstenii, Forsten’s tortoise, East Indian tortoise
    • Indotestudo travancorica, Travancore tortoise
  • Kinixys
    • Kinixys belliana, Bell's hinge-back tortoise
    • Kinixys erosa, Forest hinge-back tortoise, serrated hinge-back tortoise
    • Kinixys homeana, Home's hinge-back tortoise
    • Kinixys lobatsiana, Lobatse hinge-back tortoise
    • Kinixys natalensis, Natal hinge-back tortoise
    • Kinixys spekii, Speke's hinge-back tortoise
  • Malacochersus
    • Malacochersus tornieri, pancake tortoise
  • Psammobates
    • Psammobates geometricus, geometric tortoise
    • Psammobates oculifer, serrated tent tortoise, Kalahari tent tortoise
    • Psammobates tentorius, African tent tortoise
  • Pyxis
    • Pyxis arachnoides, (Madagascan) spider tortoise
    • Pyxis planicauda, flat-backed spider tortoise, (Madagascan) flat-tailed tortoise, flat-tailed spider tortoise
  • Stigmochelys Gray, 1873
    • Stigmochelys pardalis, leopard tortoise
  • Testudo
    • Testudo graeca, Greek tortoise, spur-thighed tortoise, Moorish tortoise
    • Testudo hermanni, Hermann's tortoise
    • Testudo horsfieldii, Russian tortoise, steppe tortoise, Horsfield's tortoise, or Central Asian tortoise
    • Testudo kleinmanni, Egyptian tortoise, including Negev tortoise
    • Testudo marginata, marginated tortoise

Subfamily Xerobatinae Agassiz, L. 1857.

  • Gopherus
    • Gopherus agassizii, Mojave Desert tortoise, Agassiz's desert tortoise
    • Gopherus berlandieri, Texas tortoise, Berlandier's tortoise
    • Gopherus flavomarginatus, Bolson tortoise
    • Gopherus morafkai, Sonoran Desert tortoise, Morajak's desert tortoise
    • Gopherus polyphemus, gopher tortoise
  • Manouria
    • Manouria emys, Asian giant tortoise, brown tortoise (mountain tortoise)
    • Manouria impressa, impressed tortoise
  • Stylemys (Genus extinct)
    • Stylemys botti
    • Stylemys calaverensis
    • Stylemys canetotiana
    • Stylemys capax
    • Stylemys conspecta
    • Stylemys copei
    • Stylemys emiliae
    • Stylemys frizaciana
    • Stylemys karakolensis
    • Stylemys nebrascensis (syn. S. amphithorax)
    • Stylemys neglectus
    • Stylemys oregonensis
    • Stylemys pygmea
    • Stylemys uintensis
    • Stylemys undabuna

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